Study review by Kip Hansen
This essay is about coyotes!
One of the odd things about this blog — WUWT — is the broad range of interests of the readers here. Perhaps I shouldn’t be surprised by this as readers here tend to be better-than-average educated, well read, interested in all things Science, concerned about the environment and tend to have more open minds.
At least three times in the last couple of years, I have written about some topic, only to have the comments section overwhelmed by discussions of coyotes — their habitat, range and behaviors — with lots of interesting stories of personal sightings and experiences.
We hear and read so much news about the threat of species extinction and shrinking ranges of species that I though a modern success story was in order.
The publishing of a brand new study about North American coyotes and their historic ranges has presented this opportunity to write about coyotes and allow readers to share their stories — this time on topic!
The new study comes to us from James W. Hody (North Carolina State University) and Roland Kays (North Carolina Museum of Natural Sciences) in a paper published in the open-access journal ZooKeys, titled Mapping the expansion of coyotes (Canis latrans) across North and Central America.
The paper is a major effort exploring this statement:
“The geographic distribution of coyotes (Canis latrans) has dramatically expanded since 1900, spreading across much of North America in a period when most other mammal species have been declining. Although this considerable expansion has been well documented at the state/provincial scale, continent-wide descriptions of coyote spread have portrayed conflicting distributions for coyotes prior to the 1900s, with popularly referenced anecdotal accounts showing them restricted to the great plains, and more obscure, but data-rich accounts suggesting they ranged across the arid west.”
Hody and Kays dug into biological history using “archaeological and fossil records, museum specimens, peer-reviewed reports, and records from wildlife management agencies” to determine the true historical range of the coyote as far back as 10,000 years before the present. What they found was that “coyotes have been present in the arid west and California throughout the Holocene, well before European colonization. Their range in the late 1800s was undistinguishable from earlier periods, and matched the distribution of non-forest habitat in the region.” Here’s the primary map they offer:

While we see that there are a few outliers, it is clear that, historically, coyotes have been mainly found in grasslands and arid lands of the North American west. The authors conclude:
“These data indicate that that coyotes’ range in the late-1800s reflected a longstanding geographic distribution that formed well before the 1700s, not a recent westward expansion. This contradicts widely-cited descriptions of the historical distribution of coyotes (Figure 1), which suggest that California and the Rocky Mountains as areas that were colonized by coyotes as recently as the 19th and 20th centuries ….. Instead, the historical distribution of coyotes matches areas where non-forested habitats (e.g., grassland, prairie, desert) dominate the climax vegetation, more closely corresponding to earlier range descriptions by Nowak … and Young and Jackson …. The Holocene distribution of coyotes in Mesoamerica remains unclear due to the relatively small number of published historical specimens available from this area.”
Using contemporary reports from the literature and various state wildlife agencies, Hody and Kays construct the following map of the expansion of the coyote’s range in North America to occupy all of the contiguous United States, all of Mexico as well was expansion into much of Canada, Alaska and Central America as far south as the Panama Canal.

This extensive colonization of new territory is hypothesized to have been facilitated by a variety of circumstances:
- The extirpation of other apex-level predators throughout North America, mainly the wolf and the cougar (mountain lion) in Eastern North America and the cougar and jaguar in Central America reducing predation of coyotes by these species and increasing available prey for the coyotes.
- The conversion of forested landscapes into agricultural landscapes opening up familiar ecosystems (similar to grasslands) to the coyotes and offering new prey — farm animals such as lambs, goats, chickens etc. This is believed to be the case in North America and in Central America.
- “Hybridization of coyotes with wolves and domestic dogs in eastern North America introduced new genotypes that may have promoted colonization and survival in eastern habitats” (see the story of the “Red Wolf”). In the southeastern United States and in Central America, hybridization is primarily with domestic dog breeds. (Oddly, hybridization with wolves and dogs does not appear to be happening on the northwestern front of the coyote’s expansion.)
All-in-all, this mid-level predator is gaining territory (and genetic content) through its incredible adaptability to modern conditions and the environmental changes being made by the continued and changing human influences on landscapes.
The paper’s authors express fears of what effects the coyote may have on South American ecosystems when the coyote manages to cross the barrier currently presented by the Panama Canal and the dense forests of the Darién Gap in southern Panama and northwestern Columbia. “If coyotes reach South America, it is likely that the grassland and agricultural habitats in Colombia and Venezuela could support viable populations, unless competition with native carnivores restricts them….. its potential effects on native wildlife is entirely unknown.”
The paper is available in pdf format from the publisher.
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Author’s Comment Policy:
North American native cultures commonly contain myths and stories revolving around the coyote, where it is often portrayed as “The Trickster”. For the Navajo, “coyote is an irresponsible and trouble-making character and he is one of the most important and revered characters in Navajo mythology.”
I currently live at the foot of the Catskill Mountains in Central Hudson Valley of New York State. The Catskills comprise 1,120 square miles (716,800 acres or 290,000 hectares) of wooded hills and valleys with an average altitude of about 3,000 feet (~1000 meters). Coyotes live and breed here and are a pest species for ranchers and farmers — one of my sons hunted them for a local farmer. In the winter, the mountain population moves downslope into the Hudson Valley which is much more densely populated. It is believed that the presence of coyotes keeps down the feral cat population (a plus).
Throughout New York State, there is a long tradition of scary stories being spread about “coy-dogs” and “coy-wolves”, often used as a “boogeyman” to prevent children was straying too far from home after dark. “Don’t go too far from the house, the coy-dogs’ll get ya!”
The American coyote is the true winner in the competition for America’s Most Successful Predator (second to Man, of course).
If you want me to respond specifically to a question or comment, address it to “Kip…” so I am sure to see it.
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Kip,
Plenty of coyotes around here in Santa Fe, NM. Some year to year variability. Haven’t noticed a trend in 12 years.
Overall they are pretty harmless and do some good in controlling rabbits and mice.
They are a pain in the butt though if you like to walk your dogs off leash. The dogs want to chase them, which is not a good idea. Occasionally they seem to follow us and lurk. One was hanging around at night below my house a few weeks ago and one of my dogs got into it with him. No real damage done but we are being a bit more cautious.
In any event, I do pack some heat in the mornings when we walk off leash.
What dd hit the press about six or so years ago was then Texas Gov Rick Perry shooting a coyote when out walking his dog. The animal rights activists were about as outraged as the gun control types.
Tom ==> Apparently the Gov’na packs a little heat when walking his dog too.
Tom,
That’s what’s great about Wyoming. No big cities to coddle idiots, though even so we have some, mostly in our bigger towns and some union bastions even in rural areas. When you need to shoot a lion or bear in town now and then, it kind of puts the damper on peta and gun control types. Seems like even here, though, the media moguls, such as they are, are all lefties. And even here people are people and with a high proportion of government employment, and the natural propensity for folks to want free shit, even our supposedly Republican government is not really very conservative with people voting for cowboy hats and pretty faces.
All in all, though, with our biggest towns only at about 40 to 50k population (and only a couple of them), little available water, total state population of around 500k (kind of like a big city suburb), and cold and windy winters, we are about 30 to 40 years “behind” the rest of the country in terms of adopting the liberal bs. But it is continually getting worse, just like Texas. We need a Rick Perry.
And, of course, all bets are off for Jackson, WY, which is a suburb of Hollywood with Cody quickly catching up. The remainder of the state is the best in the USA, and I’ve worked in all of them and have a pretty good idea of that of which I speak. People were not meant to live on top of each other. Coyotes, lions and bears are just fine.
JimG1 ==> Yeah …. I was going to counter with Jackson…..maybe they’ll secede.
Mark ==> Thanks for the first-hand from Santa Fe. Discourage your dogs from romantic liasions with the coyotes…..
While I didn’t read every post I didn’t see any mention of raccoons. In the mid-1990s we had a lot of raccoons. Fourteen in our back yard one New Year’s eve, two large adults sows, one very large boar, a bunch of very young ones, and some “teenagers.” After Lake Jackson went down and coyotes migrated into the area, few if any raccoons. Before and after the lake disappeared we had foxes. We seldom see any today but it wasn’t because of coyotes but a parvo virus epidemic. It is possible the coyotes have not allowed the fox population to recover.
Edwin ==> So many things can happen in 20 years that generalities are hard to make about animal populations. Raccoons are probe to a nasty parvovirus…
Packing up is standard coyote behavior. Always has been and always will be. Especially in winter, when they hunt larger game rather than relying on smaller prey. Same goes for partnering with badgers. They didn’t get that behavior from breeding with wolves.
Coyotes or something very like them are the ancestor of wolves.
Dogs do indeed pack up. I’ve shot lots of dogs as well as coyotes, defending my livestock, pets and family. Finally got permission to shoot some of the wolves which have been released into my AO.
The wonderful thing about imported wolves is that they wear radio collars. The state F&G maintains a Web site showing their location. Thus I can watch them watching me.
Here is a photo of some coyotes on the ice of Lake Revelstoke eying some deer that they have herded on to the ice. These guys weren’t as big as the local timber wolves but I think the male was easily 60+ pounds. Locally, in town, many cats and small dogs have been taken by coyotes and a woman was attacked and injured while defending her dog.
https://www.flickr.com/photos/85026022@N00/5760971034/in/photolist-9M5upo-9M5tYN/
E.J. Mohr ==> Excellent photo. Thanks for the coyote news from British Columbia.
Just for clarity here are the mule deer that the coyotes had herded on to the ice. Suppose if I was smart I would have combined it into the same post.
https://www.flickr.com/photos/85026022@N00/5760972460/in/photolist-9M5upo-9M5tYN/
E.J. ==> I suppose they harass the deer until one is injured or left behind because it is ill….
EJ,
Thanks!
Great pix.
Kip,
Coyotes don’t kill just the old, sick and young. When they pack up, they can kill healthy adult white tails and even mulies. One darts in and takes a bite out of a vital or sensitive area, or wherever it can. Then the pack trails the victim, taking more bites when opportunity arises.
(Reposted in correct spot.)
From what I can tell they like to herd deer on to the ice where slipping and falling can be fatal with a pack of hungry predators circling. This shot was late in the season with lots of snow on the ice making for better traction, but the new danger is breaking through the ice. Since deer are more likely to break through the ice than wolves or coyotes, the predators take advantage and hope for ready meal.
E. J. ==> Yes, that’s my opinion as well….
Bear in mind that coyotes in the same area are liable to be related or at least know each other. When two pair up, they’re often brothers. And a pack can be litter mates.
Although you can’t tell from the pictures this coyote mule deer encounter was fairly prolonged. Initially, the two coyotes were close to the deer, who were nervously looking over their shoulders in the opposite direction. The coyotes then ran towards me and the deer also moved closer. Yet you can see some of the deer locking back, which indicates to me that there was something else they were watching – likely more coyotes. Also of note was that this was midday and usually you never see coyotes or wolves since its open season on these guys.
And in the urban coyote realm we have this incident from a little over a week ago in the Vancouver Metro area – Burnaby specifically:
https://ca.news.yahoo.com/toddler-left-more-100-stitches-133444873.html
If only urban coyotes ate only cats.
But they don’t.
E. J. ==> Yes, toddler attacked by coyote just outside his backyard.
As is par for this type of thing a B.C. Conservation Service spokesperson Alicia Stark said that officials were running DNA tests to confirm it [a coyote tracked and shot] was the same animal that attacked the toddler.
Stark said the child’s injuries are unusually serious for a coyote attack.
“Coyote attacks are very rare in this area … when we do see them they are minor, maybe a scratch or a small bite.”
Yet another warning that wild animals are just that, wild — and can be dangerous.
Coyotes attacks on humans are increasing. In the 30 years leading up to March of 2006, at least 160 attacks happened in the US. While only 41 attacks happened in the almost ten years between 1988 and 1997, there were 48 attacks verified in the five years 1998 to 2003. It should be noted that the majority of these instances happened in southern California, close to the suburban areas that border wilderness.
To stalk humans, coyotes often work in teams, as they do when hunting other larger prey, attacking from the rear and flanks. Some of their successful, persistent hunts will last as long as 20 hours or so. Their unsuccessful hunts may last eight hours before they give up.
In 1981 in Glendale, California, a coyote attacked toddler Kelly Keen, who was rescued by her father, but died in surgery due to blood loss and a broken neck. In October 2009, Taylor Mitchell, a 19-year-old folk singer on tour, died from injuries sustained in an attack by a pair of coyotes while hiking the Skyline Trail of the Cape Breton Highlands National Park in Nova Scotia, Canada. It’s possible, however, that the large northeastern coyotes responsible for that particular attack may have in fact been coyote-wolf hybrids (coy-wolves), due to interbreeding with wolves when coyotes moved into eastern North America.
No doubt low-ranking wolves were delighted to make love rather than war with the invading canids.
Kip, since pictures tell the story quite nicely, here is one from earlier in the winter showing what happens when deer end up on slick ice. I didn’t see the culprits behind this particular kill, but it occurred the night before I took this photo. Some years if the ice cover is heavy you can see blood spatters and ungulate carcasses in many areas on this lake.
I’ll also add that when I was a youngster living in the Kamloops Ashcroft dry belt area it seemed the coyotes were much smaller than the ones I’ve seen of late. So, although I’m speculating, maybe there are some wolf-coyote hybrids here in western Canada. It certainly wouldn’t surprise me.
https://www.flickr.com/photos/85026022@N00/5402658763/in/dateposted-public/
E. J. ==> well, ice is a dangerous place for hooved animals — and older gents like me. At least the clean-up crew is hard at work on that unlucky deer.
Effects of coyotes’ invading the NE upon bobcat behavior, such as the felids’ switching to hares because the canids outcompete them for deer. Also, packs or pairs of coyotes killing bobcats:
http://carnivoraforum.com/topic/9396136/1/
Felix – thanks for the link to the bobcat coyote articles. I had a quick read and learned some things. I was quite interested in the average coyote weights quoted in the study in Kansas which comports nicely with the coyotes I recall seeing as a kid. Thos long ago guys were bigger than foxes, and 20 to 30 lbs seems about right vs the much larger ones that I photographed with mule deer.
You’re welcome.
IMO there were some big coyotes in the past, too.
There has also always been inbreeding with wolves, and with dogs since the first ones arrived with the Paleoamericans.
Kip – thanks very much for this article (and the link to the Red Wolfe one). I always read your articles with interest, but this was a very enjoyable change of pace. I have one question, though: you seem convinced that coyotes rarely hunt in packs?
My impression of non-urban coyotes is that in the warmer months they tend to hunt alone or in pairs (and concentrate on smaller prey), but that in the winter they commonly hunt in small packs and put in the effort to take prey up to adult deer in size (stories from friends who hunt and/or are wildlife biologists and reading the surprisingly scant literature that was available at the time [`10 years ago]). These packs may be ‘family groups’, but they are still composed of 3-7 (in my experience) adult coyote. Urban coyotes (I lived in Edmonton, AB which is supposed to be home to 3,000 coyotes) that I have seen (walking to work in the pre-dawn winter – I’d see one or two a fortnight) usually were alone, except in the river valley which is more or less bush and has deer.
Re size: Coyotes probably follow Bergmann’s Rule – populations show a positive increase in size with latitude and altitude. So, the more northern and higher elevation populations will be larger than their southern relatives with or without any hybridisation with wolves or domestic dogs. Of course, deer also follow Bergman’s Rule, so they won’t be any easier to catch, but I wonder if the southern, relatively small, coyotes are less likely to form packs and spend most of their time hunting small prey?
DaveW ==> The hunting practices of coyotes are apparently changing along with its range and possibly hybridization — with more pack/group hunting being seen and reported. I’d like to see more research on this before I formed a strong opinion. When food is scarce — winter — packing up to make large kills could be intentional (as with established wolf packs) or serendipitous — with more and more coyotes being drawn to the noise and excitment of the chase.
Coyotes or any animal for that matter, can’t take the chance of being injured when hunting their prey.
In the wild there are no hospitals, injuries can be a death sentence.
Meh. My great grandfather came over on the Oregon Trail in 1878. There were a LOT of predators that would eat you or any other prey animal as a mid morning snack. We tried to eradicate them. Didn’t work. They are in our habitats now instead of us in theirs. So learn to shoot and carry when you go for a walk..
Pamela ==> Many of our readers report doing just that….
I had to look up Coyotes in Pennsylvania (my home state) and I got a pretty good article here and I learned a lot…They are increasing in numbers, and they seem to get along with the red foxes…:in PA:
http://www.pennlive.com/wildaboutpa/2018/02/coyotes_in_pennsylvania_whats.html
There are some pretty large Coyotes in PA:
http://www.pennlive.com/pa-sportsman/2016/02/largest_coyotes_in_the_mosquit.html
J Philliup ==> Yes anything over 40 lbs is BIG for a coyote. Pennsylvania must feed them well.
J Phillip ==> Thanks for the link.
I found a coyote kill of a full sized deer a few years ago. I am in Northern Wi on the east side of the Nicolet Nat Forest in some fairly remote country. I drove down a forest road on the way out to the local pub for dinner. We had 4″ of fresh snow and the road was not plowed.
On the way home I came across the kill spot. There was a big flat area around the deer carcass where there were a million little paw prints that had packed the snow down hard. The carcass itself was stripped completely clean with a few bones missing. It was taken down and eaten clean in less than 4 hours. It wasn’t there when I went out and mine were the only tracks on the way home. My best guess is that they drove her into a fence and got a hold on her.
I was surprised to see a coyote kill on a grown deer, but it was winter and critters get hungry in the winter. I was also surprised how fast they stripped the deer down to nothing. Between that and the size of the packed snow area I would guess that there were pretty many ‘yotes involved. There was plenty of blood, so it was a fresh kill.
Ray ==> Great first-hand report of pack hunting by coyotes.
Re Coyotes in urban areas -I recall reading about a coyote which took up residence in Central Park in NYC -cant get much more urban than that.
Down under we have many instances of animals surviving very well in urban areas such as foxes living in many suburbs, while close to the CBD of Melbourne ( population near 5 million) there is a large urban park -the Fitzroy gardens- where the resident native predator is the aptly named Powerful Owl which silently swoops down on unsuspecting possums (a small mostly tree- living marsupial about the size of a small cat) which have also become widely adapted to living in urban areas
Thomho ==> No coyotes in Australia, right? Just dingos?
Kip,
I really appreciate your post on coyotes, and especially those comments relating to the east of the Mississippi River. It took me years to convince the VA DGIF that the coyote hybrids were traveling over 15 -20 miles daily/nightly and the territories of individual packs were overlapping as my guess was that they would stray more than 30 – 40 miles over time, depending on the availability of food.
Finally the DGIF has reached the same conclusion as I have. One collared female was traveling over 30 air miles between the Homestead (VA) and the Greenbrier (WV) and I believe the round trip was about 1 – 2 weeks. This was over 3 major mountains and at least 1 major river. They discovered the location of another by accident in an area east of the mountains where a collared ‘coyote’ had moved more than 60 miles away in less than 5 days and they lost track of it. It was accidentally discovered when the signal was picked up during air travel not related to that particular ‘coyote’.
The nose on a ‘coyote is extremely good. My bird dogs can pick up turkey scent over 1 1/2 miles away when the conditions are right so I’m sure a coyote can do better. Tracks in the snow show them at a full trot for 10 miles following mountain logging roads. Don’t know how far they had already traveled before I cut the tracks or how far they went after I left it. Most times there will be 2 or 3 running together and sometimes more. The wind tends to follow the valleys to a degree in the Appalachian mountains and ‘coyotes’ do the same. Another issue in favor of the coyotes is thermals. Without forced wind from atmospheric conditions the wind often dies down approaching dusk and will begin again late morning. These uphill and downhill thermals can create a lot of fairly laminar air movement, enough to blow your hat off in some cases. At around dusk a coyote can run the low route and pick up on any scent drifting down the mountain. After dawn they can run the higher route and pick up any scent drifting up. The scent identifies prey and all they have to do is literally follow their nose. A ground nesting bird has little chance in these conditions. A biologist that was educated without the full understanding of these thermals is lost with the wind. In flat country there are basically no thermals thus no effect or real life understanding. I have tried to provide education to the educated with some success, I hope.
On closing let me say that a good dog trained to strike (bark when smelling intended quarry) out of the window is an invaluable asset. I’ve had 4 of those over the past 30 years. Accuracy is near 100%. I know what can’t even be seen! A streamer on the vehicle antenna even tells me which way to go if it’s not a “hot strike”. Pheasants in SD are cheaters because when they hear the ‘dog strike’ they run a mile in the other direction so best option then was to go to the road upwind and meet ’em half way. That took a couple of days to figure out but worked out well.
Anyway, I want to know how much of the mid-Atlantic “coyote” problem was deliberate and who the players were.
“In flat country there are basically no thermals thus no effect or real life understanding”
Nonsense. Thermals occur mostly in flat country.What you are talking about isn’t thermals, it’s katabatic and anabatic winds.
tty,
Ground level thermals that would carry scent in a manner that a coyote could utilize for locating prey is very common in mountains flowing uphill and down. There is no elevation changes to facilitate this in flat country with the exception of wind drift toward the rising sun in flat land. In mountains the after dusk down draft can continue for a few hours after sunset.
Maybe you can explain the physical process that would force thermal air movement at ground level in flat country that would exceed that in a mountainous setting.
Anyway, your response is nonsense in the context of a predators utilization of the wind drifts. Spare me the technical terms stay OT with regards to the comment above. Maybe you can address the VA Trappers Association and tell them they don’t know where to set their traps. They will get a smile from it.
eyesonu ==> “I want to know how much of the mid-Atlantic “coyote” problem was deliberate and who the players were.”
Not sure I understand the question….
Kip,
No question asked. But to put my comment in a question form: Were so-called “red wolves” released in areas other than the planned Alligator National Refuge in NC? Cades Cove in the NC mountains was one reported release but they disappeared. I believe they tried it again with the same results. I think it was 15-20 each release. VA got an appearance of “coyotes” and TN got some. There was a court case filed in TN over the “red wolves” in early 1990 or so (IIRC). The land owner lost on appeal due to the commerce clause, seems people would travel across state lines to hear the “wolves” howl. What a frickin’ joke. They were probably 100 miles away by then and breeding like rats.
Do “we” now have thousands of “red wolves” in captivity? Are they being bred as “red wolves” but called coyotes when set free? Anyway, the real QUESTION is: What were “they” doing with all the puppies and the original 425 +- rejects?
A few observations from North Georgia (just north of Atlants) which I have made. Take them with a grain of salt, because I’m sure my real data runs counter to someone’s model which ‘proves’ otherwise:
Coyotes were kept out of this area by wolves, a higher-order preditor.
Wolves were extirpated by our ancestors to protect their livestock (primarily chickens).
Coyotes started moving into the area from the south, starting in the mid-1990s. They crossed North of the Chattahoochee by 2000. They are now having an effect on the local deer population, and some experts claim annual deer numbers are shrinking.
The behavior of coyotes has been evolving to adapt to human suburbs. In the wild, a mating pair has pups, and train the pups how to hunt when they are old enough. During this time, they will hunt together. This family unit is called a pack. At a certain point, the adolescent pups are ‘encouraged’ to find their own hunting grounds. This was simply to ensure enough food for everyone. In the ‘burbs, though, they have found an abundance of food. The packs are not splitting up but growing with new additions.
Coyotes love fences, particularly if a pet is confined by them. They can easily jump a five or six foot fence. Your pet dog cannot. Once inside….The only thing better are wireless fences. They can just walk in and make a snack of your pet.
They are very subject to getting rabies. They are scavenger preditors, making meals of any sick prey they may find – including those carrying rabies. There have already been attacks on children in my area by rabid coyotes.
Coyotes bring out the ignorance and stupidity of certain people who blame man for everything. “We are seeing coyotes because we are encrouching on their territories. They have a right to be here.” Nope, and nope. They are an invasive species, and who or what gave them any rights?
“Just leave them alone. They won’t hurt anyone. ” Yes, they will, and please explain why it is safe to harbor feral canines in our neighborhood, yet we routinely round-up domesticated stray canines and destroy them for our safety.
Finally, so-called experts have told us not to bother trying to get rid of them, they will just come back. So consider this: wolves were sufficiently smart and vicious to drive coyotes out. Our ancestors were sufficiently smart and vicious to drive out wolves. But now coyotes can invade the area at will? If were aren’t smart and vicious enough to drive them out, let’s bring back wolves for a few years to do it, then get rid of the wolves again!
I predict nothing will get done until the existing food sources become insufficient for their growing population. Then they will turn to a new food source; critters that weigh less than a large fawn, are much slower, and have no defenses. They are also found trapped within fences next to buildings with signs advertising daycare. I just hope our indigenous animals will still be around when that day comes.