Monarch Butterflies: a New Mystery Unfolds

Guest Essay by Kip Hansen — 17 October 2024 — 1700 words/7 minutes

The marvelous mysterious Monarch Butterfly is at it again:  baffling lepidopterists — a branch of entomology concerning the scientific study of moths and the two superfamilies of butterflies — with questions they can’t answer.

The observational evidence is this:

It is clear that [Monarch] winter colonies in Mexico are declining, yet some recent studies suggest that summer breeding populations are relatively stable and similar to historical abundances.

Here they are speaking of the Eastern United States Monarch population — the Monarch population that migrates from as far north as Canada every fall south to the mountains of central Mexico.  If your are not already familiar with the mysterious nature of this phenomenon, read my primer on the Monarch Migration or any of my earlier Monarch essays here.

There exists a modern scientific controversy about the monarchs and their migration – such a big hullabaloo that I have named it The Monarch Wars.   The basis of the Monarch Wars is:

1)  Some governmental agencies, scientists and activist groups want to declare the Monarch Butterfly, or one of its two Northern Hemisphere populations, as an Endangered Species.  In 2022, the U.S. Fish and Wildlife Service “determined that listing the monarch under the Endangered Species Act is warranted but precluded at this time by higher priority listing actions.”  The IUCN unilaterally declared the Monarch Endangered and in 2022 added it to the IUCN Red List as Endangered.  However, IUCN then delisted monarchs in 2023, declaring them only “Vulnerable”. 

2)  Other scientists, groups and interests oppose the listing of this nearly ubiquitous species as endangered – showing evidence that it is not suffering population declines or fragmented populations.

3)  There is a third view that proposes that the Monarch Migration itself, as a phenomenon, should be declared Endangered.

The evidence is unfortunately clear — for all sides in the controversy.

How is this possible?

The latest study is:

Dramatic Recent Declines in the Size of Monarch Butterfly (Danaus plexippus) Roosts During Fall Migration [ full pre-print .pdf ]

by Andrew K. Davis, Jordan R. Croy and William E. Snyder out of the University of Georgia in Athens, GA, USA.

Their Abstract starts with this line:

“The conservation status of monarch butterflies in North America is a topic of intense scrutiny and debate.”

For several years I have been calling this The Monarch Wars

Let’s look at the evidence once more:

“It is clear that winter colonies in Mexico are declining”

Clearly winter colonies in Mexico are declining – in the first 20 years of the record, there were some boom years and some low years, less than 5 hectares, but the entire two decades show much larger roosting populations.  The most recent decade has had very low roosting numbers. 

And Summer Populations?

The above image, from Crossley et al. (2022)  published in Global Change Biology  clearly shows that the summer population of Danaus plexippus, the Monarch Butterfly, seen as the vertical orange line, has not shown radical declines over the last two decades, but, if anything,  shows a slight increase of about 0.7% per year. Not a big increase, but certainly not a sharp, dangerous or worrying decline.

[Note:  The Mexico overwintering population graph covers three more years than the summer Population study.  Those three years were boom years for the overwintering population.]

The most recent study, Davis et al. (2024) is about the sizes of Monarch roosts that they form during their migration south, when they stop for the night.   Davis et al. say:

“Observations of far fewer overwintering monarch butterflies, alongside apparent rebounds during the summer breeding season, have led to heated debate whether monarchs are truly endangered. We used ~2600 citizen scientist observations of monarch “roosts” – mass aggregations of fall-migrating monarchs – to assess whether they are struggling to reach Mexico.”

And this is an important and interesting question.  Are there plenty of monarchs in the summer which then fail to reach the overwintering grounds in Mexico, and if so, for what reasons? 

Their plain language summary:

“Positive effects of a warming and greening flyway were overwhelmed by unexplained declines in roost size of up to 80%, increasing all along the path of their arduous southern migration.”

They find that “a warming and greening flyway” — and feel free to label this CO2-enhanced atmosphere and maybe climate change — had positive effects on the annual migration.  Yet there remained “unexplained declines in [in during-migration] roost size of up to 80%”, declines which increased along the southern route. 

To this author’s dismay, they conclude:

“This suggests that to save monarchs, we should focus on avoiding well-meaning efforts such as planting non-native milkweeds that foster parasites, sicken monarchs, and disrupt migration.”

I do not think that that conclusion is warranted or supported based on the evidence of the study.  It is a valid hypothesis – a possible explanation – but certainly not strongly supported.  Like all hypotheses, it is a guess, based on some data. 

They are correct when they say “This suggests that climate change might generally be benefiting monarch migration by creating milder flight conditions and enhancing nectar availability along the flyway (but see 24).”

Ah, yes, there is a but.  You can read the but here. It is a complex calculation of “energy expenditure” that might be found true of migrating monarch butterflies under warmer temperatures, not of real  butterflies actually migrating, of course. 

The reasoning of Davis et al. goes like this:

That is, while roost sizes were declining throughout the flyway, these declines grew increasingly severe further along the migration route. This latitudinal gradient in roost size declines would be consistent with increasing mortality during migration and/or monarchs increasingly abandoning migration as they move south. This apparent disruption of migration might be the missing puzzle piece that explains relatively stable summer populations in the Midwest and declining overwintering populations in Mexico.”

Alas, Davis et al., despite being a very well done investigation,  offers no definitive or actionable findings.  “Then what is driving the clear, dramatic declines in roost size seen in these data? We can only speculate,…”.

And speculate they do. While none of the speculations are new, they include:

1) Prevalence of the monarch parasite Ophryocystis elektroscirrha (“OE”) which has increased over the last decade, some say “ten-fold”.

2) The widespread planting of non-native milkweeds such as Asclepias curassavica and Calotropis gigantea by homeowners and even land managers. Some feel that this leads to more parasitisation. There have been some studies that suggested that monarch caterpillars feeding on these species might suffer stunting or weakening.   Or that the longer growing season for these species might lure the monarchs to remain north later into the Fall.

3) The raising and releasing thousands of captive-reared monarchs each year.  Some studies have implied that captive bred monarchs are weaker or have reduced navigational ability.  There are worries that interbreeding of captive and wild monarchs will harm the species as a whole.

4)  Finally, year-round resident populations of monarchs appear to be increasingly common and growing along the western and southern edges of the species range.  Vague worries about interbreeding of non-migrating monarchs and migrating monarchs and thus genetic dilution are included.  This is the “monarchs increasingly abandoning migration as they move south” hypothesis. Another possibility is that more-and-more monarchs migrate to the South, find it perfectly suitable, and settle down there for the winters.

All, as in every single one, of the monarch butterflies found in northern regions,  where winters are freezing and below, are the offspring of monarchs that have migrated north from overwintering somewhere warmer in the south.  Thus, stable or even growing summer populations in the north seems inconsistent with very small numbers of monarchs overwintering in Mexico. Yet it is the case.

West Coast Monarchs

A similar, but more profound, situation was found with the West Coast monarch population in the winters of 2020-2021 and into the next year, 2021-2022:

You see, that recovery is simply biologically impossible.  Thus a mangled version of the Sherlock Holmes quote can be applied: “When you have eliminated the impossible, whatever remains, however improbable, must be the truth”.  For that recovery, which is a physical fact, we must eliminate the impossible and doing that requires that the monarchs were overwintering somewhere – just not where they were expected and thus uncounted.

This is one of the speculative explanations for the mystery of the Eastern Population as well – the monarchs may be overwintering in Mexico after all, just not at the usual or known locations where they are expected, thus failing to be counted.

Bottom Lines:

1)  We simply don’t know how or why this situation is occurring:  far fewer Monarchs overwintering in Mexico yet stable or even growing population in the summer in the north.

2)  There are four or five good speculative answers, but none of them stand out or have stronger evidence.  It is more probable that the situation is a combination of one or more of the suggested explanations; or the dreaded something else.

3)  One thing is fairly certain now:  the slight warming seen in continental North America over the last 20 years, of maybe 1.5 degrees F, commonly called climate change, is not the cause but rather more likely a mitigating factor benefiting the monarchs.

# # # # #

Author’s Comment:

In my part of the United States, I saw very few monarchs this last summer. 

I am fascinated by the evidence about monarchs and the growing realization that so much of it driven by a “Save the Monarchs” bias.  As in almost all fields of science, the desires of researchers lead directly to their findings.  Those looking for disasters will find them, as has been the case with the monarchs.  But those researchers are stymied by other evidence – stable summer populations. 

In science, the evidence should lead to a proper framing of the situation, which, if it is deemed undesirable, then the evidence should suggest solutions that will correct or mitigate the discovered causes of the undesired situation.

Jumping the gun with solutions to problems for which the causes are only suspected lead us on a merry chase and often make the original problem even worse.

For monarchs, I still favor planting native milkweeds in your gardens (private and public), discouraging mowing of highways verges and the edges and space between corn fields, and, of course, admission that we still don’t understand what is going on with the monarchs which should lead to more and better research.

Thanks for reading.

# # # # #

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October 18, 2024 6:19 am

How well is their wintering location protected?

Is there good or any information about this species in pre colonial times? I should think this is important.

Reply to  Kip Hansen
October 18, 2024 7:48 am

The reason I asked about pre colonial conditions- is because of a debate here in Wokeachusetts. The state DFW (div. of fish and wildlife)- was promoting huge clearcuts on state forest land. I asked why. They said that there is a shortage or early succession forests and thus, a shortage of wildlife that likes that ecosystem. When most farming was abandoned here 100-130 years ago, those abandoned acreages for some decades were ideal for early succession wildlife species- like grouse, the New England cottontail rabbit, and many bird species. Often there were many old orchards too that were still productive- for the wildlife. Now, most of the forests are in the mid life range- not young stands. Sure, it’s good to have a diversity of age classes. But, in pre colonial times, those desired species were very rare- so arguing a desperate need for huge clearcuts- which of course alienate enviro groups, was a bad idea- all for species rare 300 years ago. I wanted to promote forestry work that LOOKS GOOD, that is nice thinings that left the forests looking BETTER- like parks. A half century ago, most logging here was high grading (cut the best and left the rest). I spent decades saying we need to have nicer logging- that actually is better economically, over the long term- get rid of high grading and huge clearcuts that just alienate forestry haters. I didn’t win the argument at that time- but now the enviros/greens here want to end all forestry- not just huge clearcuts, which of course emit a lot of CO2, not that I think that’s a problem. So both the state people and the enviros argue incorrectly- and I crazily got caught in the middle of these disputes.

So, regarding the Monarchs, it would be useful to have a better understanding of their population in pre colonial times- to get a sense of whether or not the species is actually endangered in any way. I wonder how much logging- good or bad- in their winter ground area effects them. Nature is complicated. Too often groups argue with slanted views but they all claim to have science on their side.

October 18, 2024 6:37 am

The ‘roosting somewhere else’ hypothesis seems reasonable.

I believe Kevin Trenberth has a new paper out, showing the missing Monarchs are hiding in the ocean.

Reply to  Pat Frank
October 18, 2024 7:48 am

Funny you say that- UFOologists now say the UFOs are hiding in the ocean. Seriously.

c1ue
Reply to  Pat Frank
October 18, 2024 9:00 am

Exactly. It seems very possible that Monarchs are simply roosting in more places, because warmer temperatures require less ideal roosting spots. So the people haunting the known “ideal” roosting spots are simply not seeing the other ones.

Reply to  Kip Hansen
October 18, 2024 4:04 pm

Kip — are there any micro GPS transponders small enough to tag a Monarch?

Jeff Alberts
Reply to  Kip Hansen
October 20, 2024 8:25 am

What about an African Monarch?

Reply to  Jeff Alberts
October 20, 2024 9:09 pm

Laden or unladen?

jshotsky
October 18, 2024 6:39 am

Monarch butterflies feed on flower nectar. Monarch butterfly larvae feed on milkweed. If there are sufficient numbers before migration, but not afterwards, they have probably not found sufficient food to survive. Warming should produce more flowers. But herbicides are widely used, which may affect the number of flowering plants along the migration route. They do migrate through my area every year, in Oregon. I’ve seen streets brown with them, and tire tracks through them in our mountains. Agriculture controls huge areas between here and Mexico. That probably accounts for declining numbers that successfully migrate. It certainly has nothing to do with Co2…

Jeff Alberts
Reply to  Kip Hansen
October 20, 2024 8:26 am

but to the coast of California south of Monterey all the way to San Luis Obispo”

At this point, that might as well be Mexico.

Scissor
Reply to  jshotsky
October 18, 2024 7:42 am

I’ve noticed more milkweed around here in Colorado than ever before. It’s rare that I see a monarch butterfly, however.

BenVincent
Reply to  jshotsky
October 18, 2024 9:49 am

Read my comment. A professor that studies Monarchs calls cities Green Deserts. Lots of green lawns but little in the way of flowers. He said Monarchs can starve before they can fly past large cities.

Reply to  BenVincent
October 18, 2024 10:26 am

They won’t starve in my back yard. 🙂

Reply to  Kip Hansen
October 19, 2024 11:59 am

This year in central New Jersey I’ve not seen many Monarchs, could be related to it being the driest summer/fall I can remember, only had to mow lawn twice!

sturmudgeon
Reply to  Kip Hansen
October 19, 2024 2:45 pm

Would they please carry away some crazy Lib judges?

Rational Keith
Reply to  BenVincent
October 18, 2024 6:35 pm

Flowers everywhere I live.

(In recent years small blue flowers and some white flowers have been spreading along sides of busy roads.)

ferdberple
October 18, 2024 6:58 am

The observer expectation bias in operation. In science you find what you expect because your expectations bias your search..

Sparta Nova 4
Reply to  ferdberple
October 18, 2024 7:38 am

And you see only what you want to see.

ferdberple
October 18, 2024 7:03 am

The only way that summer populations remain intact is that winter populations also remain intact. Thus the monarchs are simply wintering in unknown locations.

Dale Mullen
October 18, 2024 7:08 am

Kip:
As I haven’t been closely following the plight of the butterflies, the following comment is totally off the cuff and rather unscientific. However…
There has been a drastic decline in all species of butterflies during the most recent summer in Nova Scotia, Canada. With very few exceptions, the only butterfly noticed was a small white butterfly, which showed up throughout southern Nova Scotia.
No monarchs were noted and only the occasional appearance (less than half dozen) of other butterflies, other than the white species, was observed.
For whatever reason(s), we are definitely having a problem in our area.

Reply to  Dale Mullen
October 18, 2024 7:20 am

Do you have windmills around ?

Reply to  Kip Hansen
October 18, 2024 8:53 am

I can confirm the observation in so far, as in earlier times, 80ies and 90ies, after a longer car trip at late evening or night trip, I had to clean the wind shild because of a lot of dead insects and night flying butterflies. That isn’t necessary anymore.

Reply to  Kip Hansen
October 18, 2024 3:23 pm

Once I traveld 400km forth and back over weekends for a longer time. The wind shilds of the cars I used to drive didn’t change a lot in respect of aerodynamics. The most aerodynamic car I drove was an Opel GT and it was th most affected car by night insects.

ferdberple
October 18, 2024 7:09 am

40 years ago when we sailed Mexico in winter I headed south until I could sit outside at night naked and be warm.

At the time that led us to Bahia Navidad. Now according to the IPCC we should be wintering along with the monarchs in Nome Alaska.

Sparta Nova 4
Reply to  ferdberple
October 18, 2024 7:39 am

Humor – a difficult concept.

October 18, 2024 7:32 am

Thank you for keeping up with the monarch butterfly story. Last year we saw what I thought was more than usual but here on the Colorado Front Range they just aren’t very common and what I saw was unremarkable. However, this year I saw ONE. The unusual thing about that was that I saw it on my patch of showy milkweed. That patch has been there for at least 30 years, and this is the first year I recorded a single individual. I was able to photograph it and make a point to note it was there but don’t know what to think. I looked for eggs and caterpillar but found none. I did notice that it had subdued color which I found out isn’t that unusual. Nothing very scientific but certainly has me puzzled just a bit.

Reply to  Kip Hansen
October 18, 2024 10:15 am

Just throwing in my anecdotal two cents. I live in SE Virginia (Tidewater area). I plant milkweed for monarchs and parsley for black swallowtails, and this seems to have been a normal year for monarchs for me at least. I generally see one or two monarchs pretty regularly (not sure if the same one or two or different), and have seen several recently (not sure how late they’re supposed to leave this area), although I have never seen a swarm in this area.

What’s puzzling is I seem to have notice a dramatic decline in the black swallowtails. I used to plant 12 or more parsley plants so we could have some, knowing the BSTs would also eat the parsley. In the past I’ve had the parsley eaten down to bare branches by the caterpillars. The last couple years I’ve only seen a few caterpillars and a couple BST butterflies.

End of anecdotal report….

Reply to  Kip Hansen
October 18, 2024 11:19 am

Thanks for the response. At least we had more parsley for ourselves this year. 🙂

When the kids were little, I had an old aquarium tank with a screened top that I used to put the black swallowtail and monarch chrysalises in so the kids could watch them emerge and then put them on their fingers and let them go. ahh, memories…not little any more.

Reply to  Kip Hansen
October 18, 2024 1:53 pm

No grandkids yet, but momma is hoping real hard.

sturmudgeon
Reply to  Kip Hansen
October 19, 2024 2:50 pm

Gee, I thought “if you build it, they will come”.

Reply to  Kip Hansen
October 18, 2024 2:09 pm

Here is the photo. I have a video, but I don’t think I can attach it. This was taken on August 7, 2024.

August72024monarch
Reply to  Kip Hansen
October 18, 2024 4:00 pm

Well, that’s cool! Notice the subdued color?

Sparta Nova 4
October 18, 2024 7:37 am

The greens want stasis – nothing changes.
Impossible, of course.

Then there are the politicians who run on a platform of “We need change. Trust me.”

Seems greens endorsing politicians is a best a conflict of interest.

Dave Burton
October 18, 2024 7:49 am

What a fascinating article! Great job, Kip!!

JBP
October 18, 2024 8:49 am

So to summarize:

a: professional grifters have found yet another ‘current thing’ that cannot be definitively assessed or explained: so:

b. Said thing is morphed into a narrative that allows for the generation of fear-porn pushed by government, loathsome academia and media and thereafter:

c: ‘scientists’ and eco-warriors can bandwagon together to steal money (government assisted theft) to:

D. ‘stop the bad thing from happening’. Whilst:

E. Countless ‘studies’ are funded through grift to ensure the effort is never going away

then,

rinse, lather, repeat for the next current thing

Reply to  JBP
October 19, 2024 9:33 am

rinse, lather, repeat for the next current thing”

Did you mean, “Spin, blather, rebleat”? 😎

BenVincent
October 18, 2024 9:47 am

In 2022 I contacted a professor at Texas A&M that studies Monarchs. I had some questions as to whether more densely packed housing and higher 6 ft privacy fences compared to 4 ft chain link fences were a problem for butterflies. He said that butterflies fly high enough that this was not a problem.
He then went on to explain what was a problem. Lack of food and water for butterflies in large cities. Few people grow flower beds and wild flowers and plants have been replaced by lawns so there is limited food for migrating Monarchs. Monarchs also become fatigued and/or are starve before they can travel through miles of concrete and asphalt.
He calls cities Green Deserts.

BenVincent
Reply to  Kip Hansen
October 20, 2024 5:50 pm

Dr. Craig Wilson. Sorry for my delay in replying. I was busy yesterday. I went back and reread Dr. Wilson’s email to me. I misquoted him regarding green deserts. His exact words were, “Encouraging civilians to plant natives and reduce lawns that are ‘green deserts’ is a great idea but needs a whole lot of re-education.”

John Hultquist
October 18, 2024 10:20 am

Rarely do I see a large butterfly – Dry lands of Central Washington
Here are photos of what I do see:
https://northwestbutterflies.blogspot.com/2019/09/late-season-butterflies.html
Home page: Northwest Butterflies
The site is by a migrant from Kittitas County, Caitlin LaBar, now living in SW Washington State. 

Jeff Alberts
Reply to  John Hultquist
October 20, 2024 8:29 am

Don’t see a lot of butterflies on Whidbey Island either.

Bob
October 18, 2024 11:13 am

Is anyone looking for dead Monarchs along the migratory path?

sturmudgeon
Reply to  Kip Hansen
October 19, 2024 2:55 pm

Kip… I am sure you didn’t mean it in that ‘sicko’ vein. “a memorable treat) lol

Editor
October 18, 2024 2:16 pm

If there are too many to justify “endangered”, split into subspecies until there aren’t. No study needed.

Rational Keith
October 18, 2024 6:26 pm

Mebbe the sneaky critters found new hiding places in Mexico after episodes of over-population.

(Herring fish switch rivers when numbers headed for their birth estuary are high, salmon switch rivers when conditions are poor for a few weeks (they use water temperature as a measure of depth of water in path upstream to spawning areas.)

My understanding is that populations always fluctuate, weather on the migration route being a major cause.

I suggest that overall humans caused large populations by tilling ground thus helping weeds grow, new weed control methods returned populations toward historical levels.

BTW, what about the populations in the western flyway?

Fishlaw
October 18, 2024 10:34 pm

Seems obvious to me: IT’S THE CHEMTRAILS!!!

Mikehig
October 19, 2024 2:39 am

Kip; Thanks for an excellent, interesting article. I was intrigued by the butterfly pics as we have one of them – vanessa atalanta – here in the UK, known as the Red Admiral. Another – vanessa annabella – looks identical to our vanessa cardui, the Painted Lady. How did the Red Admiral come to be found on both continents….transported with plants, perhaps?

The discussion in comments about abandoned farmland, succession forests, etc brought to mind an excellent book: “Wilding” by Isabella Tree. It describes how she and her husband went about re-wilding their family farm and estate in southern England. The results were extraordinary in terms of returning species, improved biodiversity and so on. It is a very heartening story and a good antidote to the endless doom and gloom. The Knepp estate is now a very popular destination for birders and general wildlife enthusiasts.
I expect you know the book but I thought it would be of interest to readers here.
There is a follow-up: “The Guide to Re-wilding” which is what it says. Even though most of it is aimed at farmers, landowners and govt agencies, it is still a fascinating read. It also includes sections addressing small areas and gardens with much useful info and advice for those who want to encourage nature in their surroundings.

Sparta Nova 4
Reply to  Kip Hansen
October 21, 2024 9:06 am

Too competitive with solar farms means difficult to impossible to accomplish.

Yooper
October 19, 2024 4:42 am

This whole discussion brings to mind the old movie “Fly Away Home”. Somehow someone needs to track the migration to see if there are really new overwintering locations.

beanleft
October 19, 2024 5:14 am

Not knowing anything about the field of lepidopterology, my first thought was maybe the data are incomplete or poor quality, thus making any conclusions or hypotheses suspect. I think accurately counting moving butterflies over such vast areas would be very difficult .

Jeff Alberts
October 19, 2024 8:51 pm

“Blasting, billowing, bursting forth with the power of 10 billion butterfly sneezes…”

Meisha
October 20, 2024 8:02 am

While Monarchs are beautiful creatures and it is enjoyable to see a field of them flying around (I summer in the Adirondacks), I would prefer they exist to not exist. However, I cannot support the notion that all, or even any, species must be “protected” to avoid their extinction, whether humankind is the primary cause of that or not.

Nature evolves. Species come and go and “nature” moves on. For humankind to have an ethos that “nature” must stay the way it is now because we like it that way is hubris and nonsense.

Despite what some “scientists” might say, it is highly unlikely, short of global nuclear war, that mankind — at least in current times and for any foreseeable future — could somehow cause a “mass extinction” is nonsense beyond comprehension. Even short of “mass extinction,” it is highly unlikely that humankind could cause species extinction that would be detrimental to our stability as a species. Could we destroy specific experiences of beauty; e.g., seeing Monarchs? Sure, we’ve done that on local scales before (e.g., New Zealand). But on a global scale, due to humankind impact on climate (to the extent there is any)? Seriously, get a grip.

I’m not for saving species when the cost to society is more than trivial…and it should be voluntary, anyway. Using government force to require us to spend our wealth on saving species is an improper use of force, despite what some might like to force others to do or not do.

October 20, 2024 12:39 pm

We used ~2600 citizen scientist observations of monarch “roosts” – mass aggregations of fall-migrating monarchs – to assess whether they are struggling to reach Mexico.”

Along the Eastern migration route that works out to 1.15 citizen scientists per linear mile.
The Western migration route should work out to a near equal distance.

If we divvy the citizen scientists into a total 6,000 miles that works out into 2.3 citizen scientists per linear mile.

Why linear miles?
Because that is the distance, not the area.
Both Western and Eastern monarch migration routes cover immense amounts of square miles.

That paltry 2600 citizen scientists barely scratch the surface of studying monarchs, especially if their studying monarchs involves mostly driving, then binoculars.

Here in Virginia I’ve seen a few monarchs this year, not a lot. We’re mostly rural here, but suburbia has started encroaching.
I only have native milkweeds on my property.
I do have a couple of butterfly bushes, but they’re mostly visited by swallowtails and cool bee imitating hovering moths.